Temporal and Hierarchical Scales Mediate Environmental and Ecological Variability in Food Webs

Temporal changes in the environmental conditions upon which life depend are ubiquitous in nature, acting at every level of organisation from cells to ecosystems. Although the actions themselves are often poorly understood, they strongly depend on the temporal and hierarchical (organisational) scales at which they are measured; ecosystems are relatively stable through time while their species composition may undergo vast changes. Likewise any hierarchical level may be relatively stable in the short-term, but undergo vast long-term changes. This thesis aims to better understand the importance of these scales for mediating the impact of environmental variability on ecological systems.